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Gorshkov V.G. (1985) Natural selection of communities and the stability of biogeochemical cycles. In: J. Mlikovsky and V.J.A. Novak (Eds.) Evolution and Morphogenesis, pp. 787-794. Academia, Praha. Full text (PDF, 240 Kb).
The closed matter cycles should be maintained by natural selection of local ecosystems of finite size. The existence of local ecosystmes is possible provided that (1) the physical diffusion of biogenic fluxes does not exceed bioproductivity and (2) there is a definite distribution of biodestructivity over the body sizes of heterotrophic organisms.