Biotic Regulation of the Environment: Dataset for A. Makarieva's paper in Heredity 2001
Main page in English Russian version of current document Biotic Regulation of the Environment Welcome to discussion of biotic regulation!

Biotic Regulation has moved.
You are on the first web site of Biotic Regulation.
Since February 14, 2008 it is no longer updated.
Welcome to!
New address of this page: ...

Dataset for the paper by A.M. Makarieva (2001) "Variance of protein heterozygosity in different species of mammals with respect to the number of loci studied", Heredity, 87(1), 41-51. Abstract, full text (PDF, 0.4 Mb).



Protein heterozygosity of natural species of mammals

1) L is the number of studied loci, N is the number of studied individuals, HL is the expected heterozygosity.

2) Only values of the expected heterozygosity HL were considered. If it was not known from the source of data what heterozygosity value (observed or expected) was published, HL was calculated from corresponding allele frequencies, see formulas (1)-(2) in the paper. Such values are marked with an asterisk *. In case of Hardy-Weinberg equilibrium the published values were considered as values of expected heterozygosity and were included into the analysis.

3) If a certain species was represented by two or more populations where the same number of L loci was studied, the population with the largest number of studied individuals was taken. For those populations for which data for several years were available, heterozygosity values were averaged over the years and weighted by numbers of individuals studied each year.

4) If for a given species there were several values of heterozygosity obtained in different populations with different numbers of studied loci L, all these values were used only if they all belonged to different intervals of L, see Fig. 1 in the paper. If some of them fell within the same int erval, only one value of HL with the largest L was considered.

5) Only populations with ten or more studied individuals were considered.

6) In most cases data on zoo and laboratory populations of mammals were not considered.

If any problems with the presented data arise (incorrect citation, missing reference etc.), please, do not hesitate to contact the author:

  Theoretical Physics Division
Petersburg Nuclear Physics Institute
188300, Gatchina, St.-Petersburg, Russia
fax +7-813-7131963
e-mail contact:


Main page in English Russian version of current document   25 October 2001.